Abstract: This study is an in-depth extension of the key processes based on the empirical evidence of spatial genetic connotations and mechanisms of action. Following the technical path of ‘scene-feature-gene’, this paper reversely decodes the space genome associated with characteristics, focuses on the two key processes and mechanisms of space genes:hierarchical expression and intergenerational evolution, explores the intergenerational division of space genes and their characteristic factor from the common and ephemeral dimensions, and proposes the dual characteristics, paths, and mechanisms of evolution, as well as the dual mechanisms affecting the direction of gene evolution. This will provide theoretical and technical support for the inheritance and protection of characteristic genes, as well as the maintenance of the characteristics of villages and towns in multiple ways.